Publications in refereed journals
37. Walsh, J.C., Connors, K., Hertz, E., Kehoe, L., Martin, T.G., Connors, B., Bradford, M.J., Freshwater, C., Frid, A., Halverson, J., Moore, J.W., Price, M.H.H. & Reynolds, J.D (2020). Prioritizing conservation actions for Pacific salmon in Canada. J. Appl. Ecol., https://doi.org/10.1111/1365-2664.13646. Link
36. Olson, A., Frid, A., JBQ, dos S. & Juanes, F. (2020). Trophic position scales positively with body size within but not among four species of rocky reef predators. Mar. Ecol. Prog. Ser., 640, 189–200. DOI: https://doi.org/10.3354/meps13275. PDF.
35. Connors, B., Atlas, W., Melymick, C., Moody, M., Moody, J. & Frid, A. (2019). Conservation Risk and Uncertainty in Recovery Prospects for a Collapsed and Culturally Important Salmon Population in a Mixed-Stock Fishery. Mar. Coast. Fish., 11, 423–436. Link.
34. Frid, A., McGreer, M. and Frid, T. (2019) ‘Chasing the light: Positive bias in camera-based surveys of groundfish examined as risk-foraging trade-offs, Biological Conservation, 231, pp. 133–138. doi: https://doi.org/10.1016/j.biocon.2019.01.011. Link. PDF. Blog.
34.b Frid, A., Kobluk, H. & McGreer, M. (2020). Addendum to “Chasing the light: Positive bias in camera-based surveys of groundfish examined as risk-foraging trade-offs” Biological Conservation, 231, 133–138. Biol. Conserv., 244, 108513. Link. PDF.
33. Ban, N.C., Frid, A., Reid, M., Edgar, B., Shaw, D. & Siwallace, P. (2018). Incorporate Indigenous perspectives for impactful research and effective management. Nat. Ecol. Evol. https://doi.org/10.1038/s41559-018-0706-0. Link
32. Frid A, McGreer M, Gale KSP, et al. (2018). The area‐heterogeneity tradeoff applied to spatial protection of rockfish (Sebastes spp.) species richness. Conservation Letters. e12589. https://doi.org/10.1111/conl.12589. Link
31. Ban NC & Frid A. (2018). Indigenous peoples’ rights and marine protected areas. Marine Policy 87 (2018) 180–185 .Link. PDF.
30. Ban NC,Eckert LE, McGreer M, & Frid A. (2017). Indigenous knowledge as data for modern fishery management: A case study of Dungeness crab in Pacific Canada. Ecosystem Health and Sustainability. 3(8), 1379887. https://doi.org/10.1080/20964129.2017.1379887. Link. Main Points.
29. Eckert LE, Ban NC, Frid A, McGreer M. (2017) Diving back in time: Extending historical baselines for yelloweye rockfish with indigenous knowledge. Aquatic Conserv: Mar Freshw Ecosyst. 2017;1–9. https://doi.org/10.1002/aqc.2834. Link. PDF. Main Points.
28. McGreer M & Frid A. (2017). Declining size and age of rockfishes (Sebastes spp.) inherent to Indigenous cultures of Pacific Canada. Ocean and Coastal Management: 145, 14–20. Link. PDF. Main Points. Radio interview
27. Frid A., McGreer M., Haggarty D.R., Beaumont J. & Gregr E.J. (2016). Rockfish size and age: The crossroads of spatial protection, central place fisheries and indigenous rights. Global Ecology and Conservation, 8, 170-182. LINK
26. Frid A., McGreer M. & Stevenson A. (2016). Rapid recovery of Dungeness crab within spatial fishery closures declared under indigenous law in British Columbia. Global Ecology and Conservation, 6, 48-57. LINK
25. Wirsing A.J., Heithaus M.R. & Frid A. (2014). Cross-fertilizing aquatic and terrestrial research to understand predator risk effects. Wiley Interdisciplinary Reviews: Water. Published online: 4 AUG 2014. DOI: 10.1002/wat2.1039. PDF. Link.
24. Marliave J, Frid A, Welch D and Porter A. (2013.) Home site fidelity by Black Rockfish (Sebastes melanops) reintroduced to a fjord environment. Canadian Field Naturalist 127(3): 255–261. PDF
23. Frid A., Connors B, Cooper AB, and Marliave J (2013). Size-structured abundance relationships between upper- and mid-trophic level predators in temperate rocky reefs. Ethology Ecology & Evolution 25: 253-268. PDF
22. Frid A, Marliave J, and Heithaus MR (2012). Interspecific variation in life history relates to antipredator decisions by marine mesopredators on temperate reefs. PLoS ONE 7(6): e40083. PDF. LINK. Narrative version.
21. Frid A and L Quarmby (2012) Take direct action on climate inaction. Nature. 487 (5 July 2012). PDF
20. Frid A and Marliave J (2010) Predatory fishes affect trophic cascades and apparent competition in temperate reefs. Biology Letters 6:533-536. PDF
19. Frid A, Burns J, Baker GG and Thorne RE (2009). Predicting synergistic effects of resources and predators on foraging decisions by juvenile Steller sea lions. Oecologia. 158:775-7 76. PDF; Supplementary Material
18. Frid A, Baker GG and Dill LM (2008). Do shark declines create fear-released systems? Oikos 117: 191-201. PDF
17. Heithaus MR, Frid A, Wirsing, A and Worm B (2008). Predicting ecological consequences of marine top predator declines. Trends in Ecology and Evolution 23:202-210 PDF; Erratum
16. Heithaus MR, Wirsing A, Frid A and Dill LM (2008). Seascapes of fear: evaluating sublethal predator effects experienced and generated by marine mammals. Marine Mammal Science 24:1-15. PDF
15. Frid A, Dill LM, Thorne RE, and Blundell GM (2007a). Inferring prey perception of relative danger in large-scale marine systems. Evolutionary Ecology Research. 9: 635-649. PDF; Supplementary Material
14. Frid A, Heithaus MR, and Dill LM (2007b). Dangerous dive cycles and the proverbial ostrich. Oikos: 116: 893-902. PDF
13. Heithaus MR, Frid A, Wirsing A, Dill LM, Fourqurean J, Burkholder D, Thomson J, Bejder L (2007). State-dependent risk-taking by green sea turtles mediates top-down effects of tiger shark intimidation in a marine ecosystem. Journal of Animal Ecology. 76: 87-844. PDF
12. Heithaus MR, Wirsing A, Frid A, and Dill LM (2007). Behavioral indicators in conservation biology. Israel Journal of Ecology and Evolution. 53: 355-370. PDF
11. Frid A, Baker GG and Dill LM (2006). Do resource declines increase predation rates on North Pacific harbor seals? A behavior-based plausibility model. Marine Ecology Progress Series. 312: 265-275. PDF; Supplementary material.
10. Heithaus, M.R., Frid, A, Wirsing, A., Bejder, L., and Dill, L.M. (2005). Biology of sea turtles under risk from tiger sharks at a foraging ground. Marine Ecology Progress Series. 288:285-294. PDF
9. Frid A (2003) Dall’s sheep responses to overflights by helicopter and fixed-wing aircraft. Biological Conservation. 110: 387-399. PDF
8. Heithaus MR and Frid A (2003) Optimal diving under the risk of predation. Journal of Theoretical Biology. 223:79-92. PDF
7. Frid A and Dill LM (2002) Human-caused disturbance stimuli as a form of predation risk. Conservation Ecology. 6:http://www.consecol.org/Journal/vol6/iss1/art11/print.pdf.
6. Heithaus MR, Frid A and Dill LM (2002) Shark-inflicted injury frequencies, escape ability, and habitat use of green and loggerhead turtles. Marine Biology. 140:229–236. PDF
5. Heithaus MR, McLash JJ, Frid A, Dill LM and Marshall GJ (2002) Novel insights into green sea turtle behaviour using animal-borne video cameras. Journal of the Marine Biological Association of the United Kingdom. 82:1-2. PDF
4. Frid A (2001) Habitat use by endangered huemul (Hippocamelus bisulcus): cattle, snow, and the problem of multiple causes. Biological Conservation. 100:261-267. PDF
3. Frid A (1999) Huemul (Hippocamelus bisulcus) sociality at a periglacial site: sexual aggregation and habitat effects on group size. Canadian Journal of Zoology. 77:1083-1091. PDF
2. Frid A (1997) Vigilance by female Dall’s sheep: interactions between predation risk factors. Animal Behaviour. 53:799-808. PDF
1. Frid A (1994) Observations on habitat use and social organisation of a huemul (Hippocamelus bisulcus) coastal population in Chile. Biological Conservation. 67:13-19. PDF
37. Walsh, J.C., Connors, K., Hertz, E., Kehoe, L., Martin, T.G., Connors, B., Bradford, M.J., Freshwater, C., Frid, A., Halverson, J., Moore, J.W., Price, M.H.H. & Reynolds, J.D (2020). Prioritizing conservation actions for Pacific salmon in Canada. J. Appl. Ecol., https://doi.org/10.1111/1365-2664.13646. Link
36. Olson, A., Frid, A., JBQ, dos S. & Juanes, F. (2020). Trophic position scales positively with body size within but not among four species of rocky reef predators. Mar. Ecol. Prog. Ser., 640, 189–200. DOI: https://doi.org/10.3354/meps13275. PDF.
35. Connors, B., Atlas, W., Melymick, C., Moody, M., Moody, J. & Frid, A. (2019). Conservation Risk and Uncertainty in Recovery Prospects for a Collapsed and Culturally Important Salmon Population in a Mixed-Stock Fishery. Mar. Coast. Fish., 11, 423–436. Link.
34. Frid, A., McGreer, M. and Frid, T. (2019) ‘Chasing the light: Positive bias in camera-based surveys of groundfish examined as risk-foraging trade-offs, Biological Conservation, 231, pp. 133–138. doi: https://doi.org/10.1016/j.biocon.2019.01.011. Link. PDF. Blog.
34.b Frid, A., Kobluk, H. & McGreer, M. (2020). Addendum to “Chasing the light: Positive bias in camera-based surveys of groundfish examined as risk-foraging trade-offs” Biological Conservation, 231, 133–138. Biol. Conserv., 244, 108513. Link. PDF.
33. Ban, N.C., Frid, A., Reid, M., Edgar, B., Shaw, D. & Siwallace, P. (2018). Incorporate Indigenous perspectives for impactful research and effective management. Nat. Ecol. Evol. https://doi.org/10.1038/s41559-018-0706-0. Link
32. Frid A, McGreer M, Gale KSP, et al. (2018). The area‐heterogeneity tradeoff applied to spatial protection of rockfish (Sebastes spp.) species richness. Conservation Letters. e12589. https://doi.org/10.1111/conl.12589. Link
31. Ban NC & Frid A. (2018). Indigenous peoples’ rights and marine protected areas. Marine Policy 87 (2018) 180–185 .Link. PDF.
30. Ban NC,Eckert LE, McGreer M, & Frid A. (2017). Indigenous knowledge as data for modern fishery management: A case study of Dungeness crab in Pacific Canada. Ecosystem Health and Sustainability. 3(8), 1379887. https://doi.org/10.1080/20964129.2017.1379887. Link. Main Points.
29. Eckert LE, Ban NC, Frid A, McGreer M. (2017) Diving back in time: Extending historical baselines for yelloweye rockfish with indigenous knowledge. Aquatic Conserv: Mar Freshw Ecosyst. 2017;1–9. https://doi.org/10.1002/aqc.2834. Link. PDF. Main Points.
28. McGreer M & Frid A. (2017). Declining size and age of rockfishes (Sebastes spp.) inherent to Indigenous cultures of Pacific Canada. Ocean and Coastal Management: 145, 14–20. Link. PDF. Main Points. Radio interview
27. Frid A., McGreer M., Haggarty D.R., Beaumont J. & Gregr E.J. (2016). Rockfish size and age: The crossroads of spatial protection, central place fisheries and indigenous rights. Global Ecology and Conservation, 8, 170-182. LINK
26. Frid A., McGreer M. & Stevenson A. (2016). Rapid recovery of Dungeness crab within spatial fishery closures declared under indigenous law in British Columbia. Global Ecology and Conservation, 6, 48-57. LINK
25. Wirsing A.J., Heithaus M.R. & Frid A. (2014). Cross-fertilizing aquatic and terrestrial research to understand predator risk effects. Wiley Interdisciplinary Reviews: Water. Published online: 4 AUG 2014. DOI: 10.1002/wat2.1039. PDF. Link.
24. Marliave J, Frid A, Welch D and Porter A. (2013.) Home site fidelity by Black Rockfish (Sebastes melanops) reintroduced to a fjord environment. Canadian Field Naturalist 127(3): 255–261. PDF
23. Frid A., Connors B, Cooper AB, and Marliave J (2013). Size-structured abundance relationships between upper- and mid-trophic level predators in temperate rocky reefs. Ethology Ecology & Evolution 25: 253-268. PDF
22. Frid A, Marliave J, and Heithaus MR (2012). Interspecific variation in life history relates to antipredator decisions by marine mesopredators on temperate reefs. PLoS ONE 7(6): e40083. PDF. LINK. Narrative version.
21. Frid A and L Quarmby (2012) Take direct action on climate inaction. Nature. 487 (5 July 2012). PDF
20. Frid A and Marliave J (2010) Predatory fishes affect trophic cascades and apparent competition in temperate reefs. Biology Letters 6:533-536. PDF
19. Frid A, Burns J, Baker GG and Thorne RE (2009). Predicting synergistic effects of resources and predators on foraging decisions by juvenile Steller sea lions. Oecologia. 158:775-7 76. PDF; Supplementary Material
18. Frid A, Baker GG and Dill LM (2008). Do shark declines create fear-released systems? Oikos 117: 191-201. PDF
17. Heithaus MR, Frid A, Wirsing, A and Worm B (2008). Predicting ecological consequences of marine top predator declines. Trends in Ecology and Evolution 23:202-210 PDF; Erratum
16. Heithaus MR, Wirsing A, Frid A and Dill LM (2008). Seascapes of fear: evaluating sublethal predator effects experienced and generated by marine mammals. Marine Mammal Science 24:1-15. PDF
15. Frid A, Dill LM, Thorne RE, and Blundell GM (2007a). Inferring prey perception of relative danger in large-scale marine systems. Evolutionary Ecology Research. 9: 635-649. PDF; Supplementary Material
14. Frid A, Heithaus MR, and Dill LM (2007b). Dangerous dive cycles and the proverbial ostrich. Oikos: 116: 893-902. PDF
13. Heithaus MR, Frid A, Wirsing A, Dill LM, Fourqurean J, Burkholder D, Thomson J, Bejder L (2007). State-dependent risk-taking by green sea turtles mediates top-down effects of tiger shark intimidation in a marine ecosystem. Journal of Animal Ecology. 76: 87-844. PDF
12. Heithaus MR, Wirsing A, Frid A, and Dill LM (2007). Behavioral indicators in conservation biology. Israel Journal of Ecology and Evolution. 53: 355-370. PDF
11. Frid A, Baker GG and Dill LM (2006). Do resource declines increase predation rates on North Pacific harbor seals? A behavior-based plausibility model. Marine Ecology Progress Series. 312: 265-275. PDF; Supplementary material.
10. Heithaus, M.R., Frid, A, Wirsing, A., Bejder, L., and Dill, L.M. (2005). Biology of sea turtles under risk from tiger sharks at a foraging ground. Marine Ecology Progress Series. 288:285-294. PDF
9. Frid A (2003) Dall’s sheep responses to overflights by helicopter and fixed-wing aircraft. Biological Conservation. 110: 387-399. PDF
8. Heithaus MR and Frid A (2003) Optimal diving under the risk of predation. Journal of Theoretical Biology. 223:79-92. PDF
7. Frid A and Dill LM (2002) Human-caused disturbance stimuli as a form of predation risk. Conservation Ecology. 6:http://www.consecol.org/Journal/vol6/iss1/art11/print.pdf.
6. Heithaus MR, Frid A and Dill LM (2002) Shark-inflicted injury frequencies, escape ability, and habitat use of green and loggerhead turtles. Marine Biology. 140:229–236. PDF
5. Heithaus MR, McLash JJ, Frid A, Dill LM and Marshall GJ (2002) Novel insights into green sea turtle behaviour using animal-borne video cameras. Journal of the Marine Biological Association of the United Kingdom. 82:1-2. PDF
4. Frid A (2001) Habitat use by endangered huemul (Hippocamelus bisulcus): cattle, snow, and the problem of multiple causes. Biological Conservation. 100:261-267. PDF
3. Frid A (1999) Huemul (Hippocamelus bisulcus) sociality at a periglacial site: sexual aggregation and habitat effects on group size. Canadian Journal of Zoology. 77:1083-1091. PDF
2. Frid A (1997) Vigilance by female Dall’s sheep: interactions between predation risk factors. Animal Behaviour. 53:799-808. PDF
1. Frid A (1994) Observations on habitat use and social organisation of a huemul (Hippocamelus bisulcus) coastal population in Chile. Biological Conservation. 67:13-19. PDF
Peer-reviewed book chapters
2. Frid A. and Heithaus M.R. (2010) Conservation and Anti-Predator Behavior. In:Breed M.D. and Moore J., (eds.) Encyclopedia of Animal Behavior, volume 1,pp. 366-376 Oxford: Academic Press. PDF
1. Heithaus MR, Frid A, Vaudo JJ, Worm B, and Wirsing, AJ. 2010. Unravelling the Ecological Importance of Elasmobranchs. Pages 607-633 in: JC Carrier, MR Heithaus, JA Musick (editors) Sharks and Their Relatives II: Biodiversity, Adaptive Physiology, and Conservation. CRC Press. LINK
1. Heithaus MR, Frid A, Vaudo JJ, Worm B, and Wirsing, AJ. 2010. Unravelling the Ecological Importance of Elasmobranchs. Pages 607-633 in: JC Carrier, MR Heithaus, JA Musick (editors) Sharks and Their Relatives II: Biodiversity, Adaptive Physiology, and Conservation. CRC Press. LINK
